…and what percentage was Rh-? Y-DNA I2a was frequent among them. So was to an extent G2a (Caucasus origin). R1b was invasion/replacement-based. When it comes to mtDNA, Haplogroups K and J stand out. MtDNA haplogroup V showed up at a later time.and connects the Basques, Berbers, Sami of Lapland and the Guanches of the Canary islands.