Rare blood type variant cisAB found in Neanderthals

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We found that the Siberian Neanderthals, Altai and Chagyrskaya, are both homozygous for a derived Neanderthal variant of the O allele, while the European Neanderthal, Vindija, is a heterozygote for two derived Neanderthal variants, an O variant different from Altai and Chagyrskaya, and a rare cis-AB variant. The Denisovan individual is homozygous for an ancestral variant of the O allele, similar to variants found widely in modern humans.

https://www.biorxiv.org/content/10.1101/2020.07.27.223628v1.full.pdf

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What is cisAB?

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