Was the Khvalynsk culture high in Rh(D) negative frequencies?

The Yamnaya culture originated in the Don–Volga area, and is dated 3300–2600 BC. An early regional stage of Yamnaya is labeled the Mikhaylovka culture. It was preceded by the middle Volga-based Khvalynsk culture and the Don-based Repin culture (c. 3950–3300 BC), and late pottery from these two cultures can barely be distinguished from early Yamnaya pottery. Earlier continuity from eneolithic but largely hunter-gatherer Samara culture and influences from the more agricultural Dnieper–Donets II are apparent.

The Khvalynsk culture was a Middle Copper Age (for Eastern Europe named “Eneolithic”) culture (c. 4900 – 3500 BC) of the middle Volga region, discovered at Khvalynsk on the Volga in Saratov Oblast, Russia. It was preceded by the Early Eneolithic Samara culture, from which it came, and succeeded by the Late Eneolithic, Early Yamna culture, into which it developed.

Recent genetic studies have shown that males of the Khvalynsk culture carried primarily the paternal haplogroup R1b, although a few samples of R1a, I2a2, Q1a and J has been detected. They belonged to the Western Steppe Herder (WSH) cluster, which is a mixture of Eastern Hunter-Gatherer (EHG) and Caucasian Hunter-Gatherer (CHG) ancestry. This admixture appears to have happened on the eastern Pontic–Caspian steppe around 5,000 BC.

Is Rh Negative Blood Celtic?

What comes to mind immediately are the Udmurt people of the same region.